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Evolution and the Fossil Record
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John Pojeta, Jr.
Dale A. Springer
American Geological Institute
The Paleontological Society
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About the Authors
John Pojeta, Jr. has been an active paleontologist
since 1957. He is a Scientist Emeritus with the U.S.
Geological Survey (USGS) and Research Associate
with the Department of Paleobiology, Smithsonian
Institution. He earned his B.S. degree at Capital
University, Bexley, OH, majoring in biology and
chemistry and earned his M.S. and Ph.D. degrees
from the University of Cincinnati, majoring in geol-
ogy and paleontology. In 1963, he joined the USGS,
Branch of Paleontology and Stratigraphy, where he
spent his career. His research has centered on early
Paleozoic mollusks, and has taken him to many
American states, Antarctica, Australia, Canada,
China, Czech Republic, Senegal, Sweden, United
Kingdom, and elsewhere. He has been Secretary and
President of The Paleontological Society; President
of the Paleontological Research Institution; Chief,
Branch of Paleontology and Stratigraphy, USGS;
and a member of the National Academy of Sciences
Committee on Paleontological Collecting.
Dale A. Springer is a
paleontologist and Professor
of Geosciences at Bloomsburg
University in Bloomsburg,PA. She
earned her B.A. degree at Lafayette
College, Easton, PA, her M.S. degree
at the University of Rochester, NY, and
her Ph.D. at Virginia Polytechnic Institute
and State University, Blacksburg. She was a visit-
ing faculty member at Amherst and Smith Colleges
before joining the Bloomsburg faculty in 1985.
Her major research interest lies in understanding the
factors controlling temporal and spatial changes in
fossil and modern marine invertebrate communities.
Dr. Springer has a long standing interest in geo-
science education. She has served as Chairperson of
the Paleontological SocietyÔs Education Committee,
as well as on several committees of the American
Geological Institute.
Trilobite
(Ordovician)
Credits
Front cover Ð Adapted from ÑFossils Through Time,Ò a
U.S. Geological Survey poster and photographic col-
lage of life on Earth over the past 600 million years.
Pages 10-11 Ð SharkÔs tooth, Fossil seed fern, Petrified
wood (G. James)
Inside Cover and title page Ð Ammonite fossil (G. James),
Modern coral reef (J. Pojeta, Jr.), Ferns (Adobe)
Pages 12-13 Ð Hubble image, Earthrise over moon
(NASA), Trilobite (J. Pojeta, photo: G. James)
Page ii-iii Ð Trilobite (M.L. Pojeta, photo: G. James),
Fossils (J. Pojeta, Jr.)
Pages 14-15 Ð Ammonite (G. James), Block diagram
(Springer/De Atley), Stratigraphic ranges table
(modified from Edwards and Pojeta, 1994)
Page iv-v Ð Ammonite, fossil fern (G. James)
Pages 16-17 Ð Half-life diagram (modified from Bushee
and others, 2000), Ordovician limestone and shale
(J. Pojeta)
Page vi Ð Geologic Time Scale (De Atley), Adapted
from various sources
Page 1Ð Ammonite (G. James)
Page 19 Ð Forelimb comparison (modified from Daeschler
and Shubin, 1998)
Pages 2-3 Ð Chesapecten fossils (adapted From Ward
and Blackwelder, 1975 ; Bryce Canyon (M. Miller)
Pages 20-21 Ð Comparison of bird and dinosaur skeletons
and limbs (modified from Ostrom, 1975 and 1994;
Diagram comparing skulls of reptiles to mammals
(modified from Savage and Long, 1986)
Pages 4-5 Ð Trilobite, brachiopod (J. Pojeta, photo:
G. James), Tyranosaurus rex skull (Smithsonian
Institution); Jurassic Dinosaur Footprints (modified
from Haubold, 1971), Devonian and Ordovician
trilobites (adapted from Moore, 1959)
Pages 22-23 Ð Reconstruction of the Ñwalking whale that
swimsÒ (modified from Thewissen and others, 1996),
Sequoia National Park, California (Digital Vision)
Pages 6-7 Ð Charles Darwin (1875 portrait), Silurian and
Devonian fishes (modified from Fenton and Fenton,
1958), Eocene fish fossil (G. James), Jurassic/
Cretaceous fishes (modified from Romer, 1966)
Pages 24-25 Ð Brachiopod (G. James),
Dragonfly and Amphibian Fossils
(Hemera)
Design: De Atley Design
Printing: CLB Printing
Copyright ¨2001
All rights reserved.
American Geological Institute
Alexandria, Virginia
www.agiweb.org
Page 26 Ð Nautilus (G. James)
Pages 8-9 Ð Early Jurassic mammal skeleton (modified
from Jenkins and Parrington,1976), Diversification
diagram (modified from Novacek, 1994)
Back Cover Ð Grand Canyon, Arizona
(Digital Vision)
ii
EVOLUTION AND THE FOSSIL RECORD
ISBN 0-922152-57-8
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Acknowledgments
Many persons have helped us as we assembled this report. We gratefully recognize artist Julie De Atley for
the graphic design and illustration, photographer George James, Robert E. Weems (who provided the fossil
footprints), and Julia A. Jackson, Editor. We also extend our sincerest thanks and appreciation to the
following individuals for reviewing the manuscript:
David Applegate
American Geological Institute
Patricia H. Kelley
University of North Carolina,
Wilmington
Kevin Padian
University of California, Berkeley
Mel M. Belsky
Brooklyn College, CUNY
Christopher G. Maples
Indiana University, Bloomington
Kim L. Pojeta
Smithsonian Institution
David J. Bohaska
Smithsonian Institution
Sara Marcus
University of Kansas
Linda Pojeta
Northport, New York
Alan H. Cheetham
Smithsonian Institution
James G. Mead
Smithsonian Institution
Robert W. Purdy
Smithsonian Institution
Daniel Dreyfus
Smithsonian Institution
Marcus E. Milling
American Geological Institute
Vicki Quick and her students
Marshall, VA
J.T. Dutro, Jr.
U.S. Geological Survey
Don Munich
Charlestown, IN
Bruce N. Runnegar
University of California, Los Angeles
Alan Goldstein
Falls of the Ohio State Park,
Clarksville, IN
Charles Naeser
U.S. Geological Survey
Judy Scotchmoor
University of California, Berkeley
Colin D. Sumrall
Cincinnati Museum of Natural
History and Science
Pat Holroyd
University of California, Berkeley
Norman D. Newell
American Museum of Natural History
John Keith
U.S. Geological Survey
William A. Oliver, Jr.
U.S. Geological Survey
Frank C. Whitmore, Jr.
U.S. Geological Survey
The American Geological Institute and The Paleontologial Society thank the following organizations for
supporting the production and distribution of Evolution and the Fossil Record.
Publishing Partners
Supporters
Paleontological Research Institution
Howard Hughes Medical Institute
California Science Teachers Association
University of California Museum of Paleontology
Association for Women Geoscientists
National Association of Geoscience Teachers
SEPM (Society for Sedimentary Geology)
The Society for Organic Petrology
Society of Vertebrate Paleontology
Soil Science Society of America
American Institute of Biological Sciences
Society for the Study of Evolution
Cleveland Museum of Natural History
Denver Museum of Nature and Science
Sponsors
American Association of Petroleum Geologists
American Geophysical Union
Geological Society of America
California Academy of Sciences
EVOLUTION AND THE FOSSIL RECORD iii
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